The San Francisco 49ers announced on Thursday that they have signed linebacker Philip Wheeler to a one-year deal. The team also released wide receiver Trindon Holliday and waived guard Fouimalo Fonoti and wide receiver Lance Lewis.

Wheeler (6-2, 245) was originally drafted in the third round (93rd overall) of the 2008 NFL Draft by the Indianapolis Colts. Throughout his seven-year career, he has spent time with the Colts (2008-11), Oakland Raiders (2012) and Miami Dolphins (2013-14). Wheeler has played in 108 games (60 starts) and has registered 455 tackles, 5.5 sacks, four forced fumbles and two fumble recoveries. In 2013 with the Dolphins, he led the team with a career-high 118 tackles.

A 30-year-old native of Columbus, Ga., Wheeler attended Georgia Tech, where he appeared in 51 games (38 starts). He finished his career with 248 tackles, 19 sacks, 14 passes defensed, five fumble recoveries, four interceptions and three forced fumbles for the Yellow Jackets. He earned Second-Team All-ACC honors in consecutive seasons (2006-07) and was named a Second-Team All-American in 2006.

The 49ers signed Holliday (5-5, 166) on Dec. 26, 2014. He appeared in the team's final game against the Arizona Cardinals and returned three kickoffs for 83 yards.

Fonoti (6-4, 310) originally signed with the 49ers as an undrafted free agent on May 12, 2014. He was placed on the Injured Reserve List on July 30, where he spent the entire season.

Lewis (6-2, 207) signed a two-year contract with the 49ers on Aug. 9, 2014, and was signed to the team's practice squad on Aug. 31, 2014, where he spent the entire season. He later signed a Reserve/Future contract on Dec. 30, 2014.